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Daily, lunar and seasonal cycles of natural light have been key forms of environmental variation across the Earth’s surface since the first emergence of life. They have driven the development of biological phenomena from the molecule to the ecosystem, including metabolic and physiological pathways, the behaviour of individuals, geographical patterns of adaptation and species richness, and ecosystem cycles (e.g. [1–4]). Indeed, biological systems are arguably organized foremost by light [5–7].


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